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@LAST_MATCH_START
@-

$-[0] is the offset of the start of the last successful match. $-[n] is the offset of the start of the substring matched by n-th subpattern, or undef if the subpattern did not match.

Thus after a match against $_, $& coincides with substr $_, $-[0], $+[0] - $-[0]. Similarly, $n coincides with substr $_, $-[n], $+[n] - $-[n] if $-[n] is defined, and $+ coincides with substr $_, $-[$#-], $+[$#-]. One can use $#- to find the last matched subgroup in the last successful match. Contrast with $#+, the number of subgroups in the regular expression. Compare with @+.

This array holds the offsets of the beginnings of the last successful submatches in the currently active dynamic scope. $-[0] is the offset into the string of the beginning of the entire match. The nth element of this array holds the offset of the nth submatch, so $-[1] is the offset where $1 begins, $-[2] the offset where $2 begins, and so on.

After a match against some variable $var:

$` is the same as substr($var, 0, $-[0])
$& is the same as substr($var, $-[0], $+[0] - $-[0])
$' is the same as substr($var, $+[0])
$1 is the same as substr($var, $-[1], $+[1] - $-[1])
$2 is the same as substr($var, $-[2], $+[2] - $-[2])
$3 is the same as substr $var, $-[3], $+[3] - $-[3])