$-[0] is the offset of the start of the last successful match. $-[n] is the offset of the start of the substring matched by n-th subpattern, or undef if the subpattern did not match.
Thus after a match against $_, $& coincides with substr $_, $-[0], $+[0] - $-[0]. Similarly, $n coincides with substr $_, $-[n], $+[n] - $-[n] if $-[n] is defined, and $+ coincides with substr $_, $-[$#-], $+[$#-]. One can use $#- to find the last matched subgroup in the last successful match. Contrast with $#+, the number of subgroups in the regular expression. Compare with @+.
This array holds the offsets of the beginnings of the last successful submatches in the currently active dynamic scope. $-[0] is the offset into the string of the beginning of the entire match. The nth element of this array holds the offset of the nth submatch, so $+[1] is the offset where $1 begins, $+[2] the offset where $2 begins, and so on. You can use $#- to determine how many subgroups were in the last successful match. Compare with the @+ variable.
After a match against some variable $var: